Writing and Presentations

Author Information:

Name format in publications: Adam T. Clark

Google Scholar: <link>

ORCID: 0000-0002-8843-3278 <link>

Peer-Reviewed Publications:


Adam T. Clark, Jean-François Arnoldi, Yuval Zelnik, György Barabas, Dorothee Hodapp, Canan Karakoç, Sara König, Viktoriia Radchuk, Ian Donohue, Andreas Huth, Claire Jacquet, Claire de Mazancourt, Andrea Mentges, Dorian Nothaaß, Lauren Shoemaker, Franziska Taubert, Thorsten Wiegand, Shaopeng Wang, Jonathan Chase, Michel Loreau, and Stanley Harpole. "General statistical scaling laws for stability in ecological systems." Ecology Letters. in press. <link>

DOI: 10.1111/ele.13760

Michael Crawford, Kathryn Barry, Adam T. Clark, Caroline Farrior, Jessica Hines, Emma Ladouceur, Jeremy Lichstein, Isabelle Marechaux, Felix May, Björn Reineking, Lindsay Turnbull, Christian Wirth, and Nadja Rüger. "The function-dominance correlation drives the direction and strength of biodiversity-ecosystem functioning relationships". Ecology Letters. in press.

DOI: 10.1111/ele.13776

Clare Kazanski, Jane Cowles, Salli Dymond, Adam T. Clark, Aaron David, Jacob Jungers, Amy Kendig, Charlotte Riggs, Jared Trost, and Xiaojing Wei. "Water availability modifies productivity response to biodiversity and nitrogen in long-term grassland experiments." Ecological Applications. in press. <link>

DOI: 10.1002/eap.2363


Kathryn Barry, Gabriella Pinter, Joseph Strini, Karrisa Yang, Istvan Lauko, Stefan Schnizter, Adam T. Clark, Jane Cowles, Akira Mori, Laura Williams, Peter Reich, and Alexandra Wright. "A graphical null model for scaling biodiversity-ecosystem functioning relationships." Journal of Ecology 109:1549-1560, 2021. <link>

DOI: 10.1111/1365-2745.13578

Peter Hofmann, Adam T. Clark, Petra Hoffmann, Antonis Chatzinotas, Stanley Harpole, and Susanne Dunker. "Beyond nitrogen : phosphorus Estimating the minimum niche dimensionality for resource competition between phytoplankton." Ecology Letters 24:761-771. <link>

DOI: 10.1111/ele.13695

Risto Virtanen, Adam T. Clark, Michael den Herder, and Heikki Roininen. "Dynamic effects of insect herbivory and climate on tundra shrub growth: roles of browsing and ramet age." Journal of Ecology 109:1250-1262, 2021. <blogpost> <link>

DOI: 10.1111/1365-2745.13551


Adam T. Clark, Lindsay A. Turnbull, Andrew Tredennick, Eric Allan, W. Stanley Harpole, Margaret M. Mayfield, Santiago Soliveres, Kathryn Barry, Nico Eisenhauer, Hans de Kroon, Benjamin Rosenbaum, Cameron Wagg, Alexandra Weigelt, Yanhao Feng, Christiane Roscher, and Bernhard Schmid. "Predicting species abundances in a grassland biodiversity experiment: Trade-offs between model complexity and generality." Journal of Ecology 108:774–787, 2020. <link> <pdf>

DOI: 10.1111/1365-2745.13316

Christopher Avery, William Bossert, Adam T. Clark, Glenn Ellison, and Sara Fisher Ellison. "Policy implications of models of the spread of coronavirus: Perspectives and opportunities for economists." Covid Economics 12:21-68, and NBER Working Paper. <link> <pdf> <NBER>

DOI: 10.3386/w27007

Christopher Avery, William Bossert, Adam T. Clark, Glenn Ellison, and Sara Fisher Ellison. (2020). "An economist’s guide to epidemiology models of infectious disease." Journal of Economic Perspectives, 34(4):79–104. <link>


Yuxin Chen, Yuanyuan Huang, Pascal A. Niklaus, Nadia Castro-Izaguirre, Adam T. Clark, Helge Bruelheide, Keping Ma, and Bernhard Schmid. "Directed species loss reduces community productivity in a subtropical forest biodiversity experiment". Nature Ecology and Evolution 4:550–559, 2020. <link>

DOI: 10.1038/s41559-020-1127-4

Peter Dietrich, Christiane Roscher, Adam T. Clark, Nico Eisenhauer, Bernhard Schmid, and Cameron Wagg. "Diverse plant mixtures sustain a greater arbuscular mycorrhizal fungi spore viability than monocultures after 12 years". Journal of Plant Ecology 13:478–488, 2020. <link>

DOI: 10.1093/jpe/rtaa037

Canan Karakoç, Adam T. Clark, and Antonis Chatzinotas. "Diversity and coexistence are influenced by time-dependent species interactions in a predator-prey system". Ecology Letters 23:983–993, 2020. <link> <pdf>

DOI: 10.1111/ele.13500

Clarence Lehman, Shelby Loberg, Adam T. Clark, and Daniel Schmitter. "Unifying the basic models of ecology to be more complete and easier to teach". BioScience 70:415–426, 2020. <link>

DOI: 10.1093/biosci/biaa013

Lina K. Mühlbauer, Maximilienne Schulze, W. Stanley Harpole, and Adam T. Clark. "gauseR: Simple methods for fitting Lotka-Volterra models describing Gause’s 'Struggle for Existence'." Ecology and Evolution 10:13275-13283, 2020. <link> <pdf>

DOI: 10.1002/ece3.6926


Adam T. Clark, Kathryn E. Barry, Christiane Roscher, Tina Buchmann, Michel Loreau, and W. Stanley Harpole. "How to estimate complementarity and selection effects from an incomplete sample of species". Methods in Ecology and Evolution 10:2141–2152, 2019. <link> <pdf> <sup>

DOI: 10.1111/2041-210X.13285

Adam T. Clark, Helmut Hillebrand, and W. Stanley Harpole. "Scale both confounds and informs characterization of species coexistence in empirical systems". American Naturalist 194:794–806, 2019. <link> <pdf> <sup>

DOI: 10.1086/705826

Adam T. Clark, Johannes M. H. Knops, and David Tilman. "Contingent factors explain average divergence in functional composition over 88 years of old field succession". Journal of Ecology 107:545–558, 2019. <link> <pdf> <sup>

DOI: 10.1111/1365‐2745.13070

Kathryn E. Barry, Hans de Kroon, Peter Dietrich, W. Stanley Harpole, Anna Roeder, Bernhard Schmid, Adam T. Clark, Margaret M. Mayfield, Cameron Wagg, and Christiane Roscher. "Linking local species coexistence to ecosystem functioning – a conceptual framework from ecological first principles in grassland ecosystems". Advances in Ecological Research 61:265–296, 2019. <link>

DOI: 10.1016/bs.aecr.2019.06.007

Jane Catford, Annabel Smith, Peter Wragg, Adam T. Clark, Margaret Kosmala, Jeannine Cavender-Bares, Peter Reich, and David Tilman. "Traits linked with species invasiveness and community invasibility vary with time, stage and indicator of invasion in a long-term grassland experiment". Ecology Letters 22:593–604, 2019. <link>

DOI: 10.1111/ele.13220

Forest Isbell, David Tilman, Peter Reich, and Adam T. Clark. "Deficits of biodiversity and productivity linger a century after agricultural abandonment". Nature Ecology and Evolution 3:1533–1538, 2019. <link>

DOI: 10.1038/s41559-019-1012-1


Adam T. Clark, Matteo Detto, Helene C. Muller-Landau, Stefan A. Schnitzer, S. Joseph Wright, Richard Condit, and Stephen P. Hubbell. "Functional traits of tropical trees and lianas explain spatial structure across multiple scales". Journal of Ecology 106:795–806, 2018. <pressrelease> <link> <pdf> <sup>

DOI: 10.1111/1365-2745.12804

Adam T. Clark, Clarence Lehman, and David Tilman. "Identifying Mechanisms that Structure Ecological Communities by Snapping Model Parameters to Empirically-Observed Tradeoffs". Ecology Letters 21:494–505, 2018. <link> <pdf> <sup>

DOI: 10.1111/ele.12910

Adam T. Clark and Claudia Neuhauser. "Harnessing Uncertainty to Approximate Mechanistic Models of Interspecific Interactions". Theoretical Population Biology 123:35–44, 2018. <link> <pdf> <sup>

DOI: 10.1016/j.tpb.2018.05.002

Lauren L. Sullivan*, Adam T. Clark*, David Tilman, and Allison K. Shaw (*authors contributed equally). "Mechanistically-derived dispersal kernels explain species-level patterns of recruitment and succession". Ecology 99:2415–2420, 2018. <bulletin> <pressrelease> <link> <pdf> <sup>

DOI: 10.1002/ecy.2498

Heather Tallis, PeterHawthorne, Stephen Polasky, Joseph Reid, Michael Beck, Kate Brauman, Jeffrey Bielicki, Seth Binder, Matthew Burgess, Emily Cassidy, Adam T. Clark, Joseph Fargione, Edward Game, James Gerber, Forest Isbell, Joseph Kiesecker, Robert McDonald, Marc Metian, Jennifer Molnar, Nathan Mueller, Christine O'Connell, Daniel Ovando, Max Troell, Timothy Boucher, and Brian McPeek. "An Attainable Global Vision for Conservation and Human Well-Being". Frontiers in Ecology and the Environment 16:563–570, 2018. <pressrelease> <link>

DOI: 10.1002/fee.1965


Adam T. Clark. "Old Fields". In Oxford Bibliographies in Ecology. Ed. David Gibson. New York: Oxford University Press, 2017. <link> <pdf>

DOI: 10.1093/OBO/9780199830060-0188

Richard Barnes* and Adam T. Clark* (*authors contributed equally). "65 Million Years of Change in Temperature and Topography Explain Evolutionary History in Eastern North American Plethodontid Salamanders". American Naturalist 190:E1–12, 2017. <pressrelease> <link> <pdf> <sup>

(Awarded the 2018 MCED Award for Innovative Contributions to Ecological Modelling)

DOI: 10.1086/691796

Adam T. Clark, H. Ye, Forest Isbell, Ethan R. Deyle, Jane Cowles, David Tilman, and George Sugihara. "Spatial ’convergent cross mapping’ to detect causal relationships from short time-series". Ecology 96(5) 1174–1181, 2015. <link> <pdf> <sup>

DOI: 10.1890/14-1479.1

Jacob M. Jungers, Adam T. Clark, Kevin Betts, Margaret E. Mangan, Donald L. Wyse, and Craig C. Sheaffer. "Long-Term Biomass Yield and Species Composition in Native Perennial Bioenergy Cropping Systems". Agronomy Journal 107(5):1627–1640, 2015. <link> <pdf>

DOI: 10.2134/agronj15.0014

Adam T. Clark, Jessica J. Rykken, and Brian D. Farrell. "The effects of biogeography on ant diversity and activity on the Boston Harbor Islands, Massachusetts, USA". PLOS ONE 6(11), 2011. <link> <pdf>

DOI: 10.1371/journal.pone.0028045

Software Packages:

gauseR: Functions for fitting Lotka-Volterra models to time series data, and to Gause's experiments. GitHub R package. Adam Clark, Lina Mühlbauer, Maximilienne Schulze. <link>

EDMhelper: Wrapper functions for automating EDM analyses. GitHub R package. Canan Karakoç & Adam Clark. <link>

partitionBEFsp: Methods for Calculating the Loreau & Hector 2001 BEF Partition. CRAN R package. Adam Clark. <link>

rEDM: Applications of Empirical Dynamic Modeling from Time Series. CRAN R package. Hao Ye, Adam Clark, Ethan Deyle. <link>

nondirfit: Methods for Multivariate Type-II Regression. GitHub R project. Adam Clark. <link>

multispatialCCM: Multispatial Convergent Cross Mapping. CRAN R package. Adam Clark. <link>

tilmanstability: Code for calculating stability sensu Lehman and Tilman 2000 American Naturalist 156(5). GitHub R project. Adam Clark. <link>

Other Academic Writing:

Free eBook Textbook: Clarence Lehman, Shelby Loberg, and Adam T. Clark. (2019). Quantitative Ecology: A New Unified Approach. University of Minnesota Libraries Publishing. University of Minnesota Digital Conservancy, http://hdl.handle.net/11299/204551. <link> <pdf>

Exercises and practice questions for the ebook: <link>

Adam T. Clark. Constraints and tradeoffs: Toward a predictive, mechanism-based understanding of ecological communities. Ph.D. Thesis, 2017. <link> <pdf>

HDL: hdl.handle.net/11299/191380

DOI: 10.13140/RG.2.2.16076.87687

The Graduate School Cookbook: One way to approach PhD applications in ecology <link> <pdf> <github>

This document is a collection of thoughts on the graduate school application process that I've collected over the years from friends and colleagues. If you have any comments on items to add or change, please let me know, either through email, or through the Github page.

How to find a moose: intentions in phenomenological and mechanistic models. PLOS Ecology Blog, 2016. <link>

Adam T. Clark. Ant Communities of the Boston Harbor Islands National Recreation Area. Undergraduate Honors Thesis, 2011. <pdf>

DOI: 10.13140/RG.2.2.21110.04160

Selected Presentations:


Invited Seminar, Harvard Kennedy School of Government, MA, USA, "Multidisciplinary Perspectives on COVID-19". <link>

Invited Seminar, Centre for Ecological Research, Budapest, Hungary, "The Ecotones of Minnesota: Prairie, Savanna, and Forest, and all in 2200 ha". <link>

Dept. Seminar, Universität Graz, Institut für Biologie, "Ökologische Mischregion Minnesota: Prairie, Savanne, und Wald, und alles auf 2200 ha". <link>

Invited Seminar, Masaryk University, Department of Botany and Zoology, "Chasing the Dragon, and two other methods: A general approach for measuring stability in complex ecological systems." <link>


Invited Seminar, University of Graz, "Predicting ecological changes across space and time by putting theory into practice." <ppt>

Invited Teaching Seminar, University of Graz, "Veränderung der Waldgrenze in den Alpen in Lichte des Klimawandels." <ppt>

Contributed Presentation, iDiv Annual Conference, "Leveraging scale to build empirically tractable theory and metrics of stability and coexistence." <pdf>

Invited Presentation, University of Oulu, "A more generalizable approach to scale-wise analysis of stable coexistence, or, Chasing the Dragon, and two other methods." <pdf>


Contributed Presentation, Ecological Society of Germany, Austria, and Switzerland (GfÖ) Annual Meeting, Vienna, Austria. "Spatial and temporal scales confound tests of stable coexistence, but identify underlying mechanisms."


Invited Presentation, Utrecht University, "Identifying mechanisms that structure ecological communities." <pdf>

Minnesota Math Climate Seminar, University of Minnesota, Twin Cities. "How can we define and identify stability - or persistence? - in diverse ecological communities?" <pdf>

PhD Defense, Department of Ecology, Evolution, and Behavior, University of Minnesota, Twin Cities. "Constraints and tradeoffs: Toward a predictive, mechanism-based understanding of ecological communities." <pdf> <video>

Invited Workshop, Zurich-Basel Plant Science Center Summer School, Einsiedeln, Switzerland. "Empirical Dynamic Modeling: Time Series Analysis for Predicting Future Dynamics and Testing Causal Links." <pdf>


Contributed Presentation, Ecological Society of America Annual Meeting, Baltimore, MD. "A predictive community ecology model for extrapolating from field observations to novel conditions." <pdf>

Contributed Presentation, World Conference on Natural Resource Modeling: Modeling and Sustainability of Biodiversity and Ecosystem Services, Bordeaux, France. "Predicting communities from monocultures: How to maximize desired goods and services with available ecological information." <pdf>


Contributed Presentation, American Society of Naturalists meeting on integrating pattern and process to understand biodiversity, Asilomar Conference Center, Pacific Grove, CA. "Defining ecosystems by their interactions: Using process to demystify pattern." <pdf>


Contributed Presentation, International Biogeography Society special meeting on species associations, Montreal, QC, Canada. "Defining communities by their interactions: Using process to identify pattern." <pdf>

Contributed Presentation, Ecological Society of America Annual Meeting, Minneapolis, MN. "Convergence and divergence in old field succession: Are there predictable trends in long-term dynamics?" <pdf>

IGNITE Presentation, "Succeeding with succession: Using ecological community dynamics to build low-impact biofuels." <pdf>


Contributed Poster, Long Term Ecological Research Network Meeting, Estes Park, CO. "Multiple Trajectories of old Field Succession at Cedar Creek." <pdf>

Contributed Presentation, Ecological Society of America Annual Meeting, Portland, OR. "When resident species resist displacement, invading species have ecologically distinct impacts on communities." <pdf>